1. Rubenstein School of Environment and Natural Resources University of Vermont 81 Carrigan Drive Burlington VT 05405 U.S.A.
2. Gund Institute for Environment University of Vermont 617 Main Street Burlington VT 05405 U.S.A.
3. Department of Earth and Environmental Systems The University of the South, 735 University Avenue, Sewanee TN 37375 U.S.A.
4. School of Earth and Environment Sciences The University of Queensland St Lucia QLD 4072 Australia
5. Center for Biodiversity and Conservation Science The University of Queensland St Lucia QLD 4072 Australia