1. Division of Ecology and Evolution, Research School of Biology The Australian National University Canberra ACT 0200 Australia
2. Department of Animal Ecology Netherlands Institute of Ecology (NIOO‐KNAW) Wageningen The Netherlands
3. Department of Biology Utrecht University Utrecht The Netherlands
4. Department of Animal Ecology and Physiology, Institute for Water and Wetland Research Radboud University Nijmegen The Netherlands