1. Key Laboratory of Sustainable Development of Marine Fisheries; Key Laboratory of Maricultural Organism Disease Control; Ministry of Agriculture; Yellow Sea Fisheries Research Institute; Chinese Academy of Fishery Sciences; Qingdao China
2. Laboratory for Marine Fisheries Science and Food Production Processes; Qingdao National Laboratory for Marine Science and Technology; Qingdao China
3. Fishery College; Ocean University of China; Qingdao China
4. College of Fisheries and Life Science; Shanghai Ocean University; Shanghai China