1. Department of Molecular Biology and Applied Physiology; Tohoku University School of Medicine; Sendai Miyagi 980-8575 Japan
2. Technology and Development Team for Mouse Phenotype Analysis; Japan Mouse Clinic; RIKEN BRC; Tsukuba Ibaraki 305-0074 Japan
3. Department of Pathology and Histotechnology; Tohoku University Graduate School of Medicine; Sendai Miyagi 980-8575 Japan
4. Faculty of Bioscience; Nagahama Institute of Bio-Science and Technology; Nagahama Shiga 526-0829 Japan
5. Faculty of Sports Science; Sendai University; Shibata-Gun Miyagi 989-1693 Japan