1. Key Laboratory of Tropical Biological Resources of Ministry of Education, School of Life and Pharmaceutical Sciences Hainan University Haikou 570228 China
2. Department of Biology Oberlin College Oberlin OH 44074 USA
3. National Herbarium of New South Wales Royal Botanic Gardens and Domain Trust Sydney NSW 2000 Australia
4. Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Menglun 666303 Yunnan China
5. Graduate School of Human and Environmental Studies Kyoto University Kyoto 606‐8501 Japan
6. Botanical Gardens Tohoku University Sendai 980‐0862 Japan
7. Department of Botany Smithsonian Institution Washington, DC 20013‐7012 USA