1. Department of Ecology and Evolutionary Biology; Yale University; New Haven CT 06511 U.S.A.
2. Centre of Excellence for Coral Reef Studies; James Cook University; Townsville 4811 Australia
3. Ecole Pratique des Hautes Etudes, USR 3278 EPHE-CNRS-UPVD, Labex Corail, CRIOBE; 66860 Perpignan France
4. Institut de Recherche pour le développement (IRD), UMR Entropie-Labex CORAIL; Université de Perpignan; 66000 Perpignan France
5. Depto. de Ecologia e Zoologia, Marine Macroecology and Biogeography Laboratory, CCB; Universidade Federal de Santa Catarina; Florianópolis 88040-900 Brazil