1. College of Resources and Environment Qingdao Agricultural University Qingdao China
2. Key Laboratory of Biofuels Qingdao Engineering Research Center of Biomass Resources and Environment Shandong Provincial Key Laboratory of Energy Genetics Qingdao Institute of Bioenergy and Bioprocess Technology Chinese Academy of Sciences Qingdao China
3. Institute for Evolution and Biodiversity University of Münster Münster Germany