1. Institute of Natural and Mathematical Sciences; Massey University at Albany; Private Bag 102 904; North Shore Mail Centre; Auckland New Zealand
2. New Zealand Institute for Advanced Study and Allan Wilson Centre for Molecular Ecology and Evolution; Massey University at Albany; Private Bag 102 904; North Shore Mail Centre; Auckland New Zealand
3. Max Planck Institute for Evolutionary Biology; August-Thienemann Strasse 2; 24306 Plön Germany