1. State Key Laboratory of Biocontrol, School of Life Sciences and School of Ecology Sun Yat‐sen University Guangzhou China
2. Laboratory of Geospatial Technology for the Middle and Lower Yellow River Regions College of Environment and Planning, Henan University Kaifeng China
3. Center of Plant Ecology, Core Botanical Garden Chinese Academy of Sciences Mengla China
4. Center for Biodiversity, Department of Biology Temple University Philadelphia Pennsylvania USA
5. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems South China Botanical Garden, Chinese Academy of Sciences Guangzhou China