1. Hubei Collaborative Innovation Centre for Grain Industry/Engineering Research Centre of Ecology and Agricultural Use of Wetland, Ministry of Education Yangtze University Jingzhou China
2. Hubei Key Laboratory of Food Crop Germplasm and Genetic Improvement Food Crops Institute, Hubei Academy of Agricultural Sciences Wuhan China
3. Department of Biology, Faculty of Applied Science Umm Al‐Qura University Makkah Saudi Arabia
4. Agronomy Department College of Agriculture, Assiut University Assiut Egypt