1. EGFV, Université de Bordeaux Bordeaux Sciences Agro, INRAE, ISVV Villenave d'Ornon F‐33882 France
2. CSIRO Agriculture and Food Glen Osmond SA 5064 Australia
3. Beijing Key Laboratory of Grape Science and Enology and CAS Key Laboratory of Plant Resources Institute of Botany, Chinese Academy of Sciences Beijing 100093 China