1. Department of Geography and Environmental Sciences; University of Zimbabwe; PO Box MP 167, Mount Pleasant Harare Zimbabwe
2. UPR AGIRs, Cirad; Department ES, Campus International de Baillarguet; 34398 Cedex 5 Montpellier France
3. UPR AGIRs, Cirad-RP-PCP; TREP Building, University of Zimbabwe; PO Box 167 MP, Mount Pleasant Harare Zimbabwe
4. Department of Biological Sciences; University of Zimbabwe; PO Box MP 167, Mount Pleasant Harare Zimbabwe
5. Department of Zoology and Entomology; Mammal Research Institute, University of Pretoria; Pretoria 0110 South Africa