1. State Key Laboratory of Tree Genetics and Breeding Research Institute of Forestry, Chinese Academy of Forestry Beijing 100091 China
2. State Key Laboratory of Subtropical Silviculture, College of Forestry and Biotechnology Zhejiang A & F University Hangzhou 311300 China
3. Taishan Academy of Forestry Sciences Taian 271000 China
4. State Key Laboratory of Tree Genetics and Breeding, Chinese Academy of Forestry Beijing 100091 China