1. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, Department of Ecology and Evolutionary Biology Fudan University Songhu Road 2005 Shanghai 200438 China
2. Key Laboratory of Biology and Genetic Resources of Tropical Crops, Hainan Key Laboratory for Biosafety Monitoring and Molecular Breeding in Off‐Season Reproduction Regions Institute of Tropical Bioscience and Biotechnology, CATAS Haikou 571101 China