1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization; Chengdu Institute of Biology; Chinese Academy of Sciences; P.O. Box 416 Chengdu 610041 China
2. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; Kunming 650201 China
3. University of Chinese Academy of Sciences; Beijing 100049 China
4. Department of Biology; Vietnam National Museum of Nature; Vietnam Academy of Science and Technology; 18th Hoang Quoc Viet Road Ha Noi Vietnam
5. Missouri Botanical Garden; P.O. Box 299 St. Louis MO 63166-0299 USA