1. Department of Physiology, Nagoya City University Medical School, Mizuho‐ku, Nagoya 467‐8601, Japan
2. Department of Basic Human Sciences, Waseda University School of Human Sciences, Tokorozawa 359, Japan
3. Section of Physiology, Exploratory Research for Advanced Technology, Japan Science and Technology Corporation, Hon‐komagome, Bunkyo‐ku, Tokyo 113‐0021, Japan
4. Department of Molecular Neurology, Institute of Medical Sciences, The University of Tokyo, Shirokanedai, Minato‐ku, Tokyo 108‐8639, Japan
5. Laboratory for Developmental Neurobiology, Brain Science Institute, Institute of Physical and Chemical Research (RIKEN), Wako, Saitama 351‐0106, Japan