1. Department of Nature Conservation, Tshwane University of Technology, Pretoria, South Africa
2. Museum of Vertebrate Zoology, University of California, Berkeley, CA, USA
3. Department of Geography, University of Southampton, Southampton, UK
4. Wildlife Conservation Research Unit, Department of Zoology, Oxford University, Tubney, UK
5. Institute of Zoology, Zoological Society of London, London, UK
6. Department of Ecology, Evolution and Behavior, University of Minnesota, Saint Paul, MN, USA
7. Scientific Services, SANParks, Skukuza, South Africa