1. Department of Biology, University of Antwerp (RUCA), Groenenborgerlaan 171, B‐2020 Antwerpen, Belgium. E‐mail:
2. Royal Belgian Institute of Natural Sciences, Vautierstraat 29, B‐1000 Brussel, Belgium
3. Department of Animal Biology, University of Santiago de Compostela, E‐15706 Santiago de Compostela, Spain
4. Staatliches Museum für Naturkunde, PF 30 01 54, D‐02806 Görlitz, Germany