1. Division of Biosphere Science, Graduate School of Environmental Science, Hokkaido University, Kita‐ku, Sapporo, Hokkaido, Japan
2. Port of Nagoya Public Aquarium, Minato‐ku, Nagoya, Japan
3. Department of Chemistry, Biology and Marine Sciences, Faculty of Science University of the Ryukyus 1 Senbaru, Nishihara, Okinwa, Japan
4. Usujiri Fisheries Station, Field Science Center for Northern Biosphere, Hokkaido University, Hakodate, Hokkaido, Japan