1. International Coastal Research Center, Ocean Research Institute, The University of Tokyo, Otsuchi, Iwate, Japan
2. Key Laboratory of Freshwater Fish Germplasm Resources and Biotechnology, Ministry of Agriculture of China, Yangtze River Fisheries Research Institute, Chinese Academy of Fisheries Science, Jingzhou, Hubei, China
3. Freshwater Fisheries Research Center, Chinese Academy of Fishery Sciences, Wuxi, Jiangsu, China
4. National Institute of Polar Research, Itabashi, Tokyo, Japan
5. Ocean Research Institute, The University of Tokyo, Nakano, Tokyo, Japan