1. School of Biological Sciences, University of Nottingham, University Park, Nottingham NG7 2RD, U.K.
2. Biological Sciences Division, Lancaster University, Lancaster LA1 4YQ, U.K.
3. Designated Areas and Site Branch, Research and Advisory Services Directorate, Scottish Natural Heritage, 2 Anderson Place, Edinburgh EH6 5NP, U.K.
4. Faculty of Veterinary Science, University of Liverpool, Liverpool L69 3BX, U.K.
5. Department of Biology, University of Buckingham, Buckingham MK18 1EG, U.K.