1. Biozentrum Grindel und Zoologisches Museum, Universität Hamburg, Hamburg, Germany
2. Mbarara University of Science and Technology, Mbarara, Uganda
3. Unit of Evolutionary Biology and Systematic Zoology, Institute for Biochemistry and Biology, University of Potsdam, Potsdam, Germany
4. Department of Zoology, University of Oklahoma, Norman, OK, USA