1. Department of Life Science, Tunghai University, Taichung, Taiwan, ROC
2. Endemic Species Research Institute, Jiji, Nantou, Taiwan, ROC
3. Laboratory of Wildlife Biology, Obihiro University of Agriculture and Veterinary Medicine, Obihiro, Japan
4. The University Museum, The University of Tokyo, Tokyo, Japan
5. Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, Hanoi, Vietnam
6. Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming, China
7. School of Life Sciences, Xiamen University, Xiamen, China