1. Wildlife and Emerging Diseases Section, Centre for Tropical Veterinary Medicine, Royal (Dick) School of Veterinary Studies, University of Edinburgh, Easter Bush, Roslin, Midlothian, EH25 9RG, United Kingdom
2. Tanzania National Parks, P.O. Box 3134, Arusha, Tanzania
3. Ministry of Water and Livestock Development, P.O. Box 322, Musoma, Tanzania
4. Division of Environmental and Evolutionary Biology, University of Glasgow, Glasgow G12 8QQ, United Kingdom
5. Frankfurt Zoological Society, P.O. Box 14935, Arusha, Tanzania
6. Department of Ecology, Evolution & Behavior, University of Minnesota, 1987 Upper Buford Circle, St. Paul, MN 55108, U.S.A.