1. Organismal and Evolutionary Biology Research Programme, Faculty of Biological and Environmental Sciences University of Helsinki Helsinki Finland
2. Tvärminne Zoological Station, Faculty of Biological and Environmental Sciences University of Helsinki Helsinki Finland
3. Department of Ecology and Evolution, Biophore University of Lausanne Lausanne Switzerland
4. Department of Biology, Faculty of Sciences University of Copenhagen Copenhagen Denmark
5. Zoology and Evolutionary Biology University of Regensburg Regensburg Germany