1. CNRS and Univ. de Strasbourg, UMR7178, IPHC; 23 rue Becquerel FR-67087 Strasbourg France
2. Center for International Collaboration and Advanced Studies in Primatology, Kyoto Univ. Primate Research Inst.; Kanrin 41-2, Inuyama, Aichi 484-8506 Japan
3. Centre d'Etudes Biologiques de Chizé; FR-79360 Villiers-en-Bois France
4. Inst. for Marine and Antarctic Studies, Univ. of Tasmania; Hobart 7001 Australia
5. Australian Antarctic Division, Dept of the Environment; Australian Government; Channel Highway, Kingston 7050 Australia