1. Graduate School of Agricultural Science; Tohoku University; Sendai 980-8572 Japan
2. Graduate School of Life and Environmental Science; Osaka Prefecture University; Sakai 599-8531 Japan
3. Graduate School of Agricultural and Life Sciences; The University of Tokyo; Tokyo 188-0002 Japan
4. National Institute of Agrobiological Science; Tsukuba Ibaraki 3058602 Japan