1. Department of Otolaryngology, Faculty of Medicine University of Tsukuba Tsukuba Japan
2. Department of Medical Biochemistry Tohoku University Graduate School of Medicine Sendai Japan
3. Department of Anatomy and Embryology, Faculty of Medicine University of Tsukuba Tsukuba Japan
4. Division of Medical Biochemistry Tohoku Medical Pharmaceutical University Sendai Japan
5. Division of Pathology Tohoku Medical Pharmaceutical University Sendai Japan
6. Cell and Developmental Biology University of Michigan Ann Arbor Michigan