1. Department of Life Science Pohang University of Science and Technology Pohang 37673 Korea
2. Department of Integrative Bioscience & Biotechnology Pohang University of Science and Technology Pohang 37673 Korea
3. Department of Biological Sciences Sungkyunkwan University Suwon 16419 Korea
4. Graduate School of Biological Sciences and Institute for Research Initiatives Nara Institute of Science and Technology Ikoma Nara 630‐0192 Japan