1. State Key Laboratory of Crop Genetics and Germplasm Enhancement Nanjing Agricultural University Nanjing 210095 China
2. Graduate School of Agriculture and Life Sciences University of Tokyo Yayoi, Bunkyo Tokyo 113‐8657 Japan
3. Graduate School of Life Sciences Tohoku University Katahira 2‐1‐1, Aoba‐ku Sendai 980‐8577 Japan
4. RIKEN Center for Sustainable Resource Science 1‐7‐22 Suehiro‐cho, Tsurumi‐ku Yokohama 230‐0045 Japan
5. Division of Biological Science Graduate School of Bioagricultural Sciences Nagoya University Furo‐cho, Chikusa‐ku Nagoya 464‐8601 Japan