1. Division of Gene Regulation Institute for Advanced Medical Research School of Medicine Keio University Tokyo Japan
2. Department of Surgery School of Medicine Keio University Tokyo Japan
3. Department of Obstetrics and Gynecology School of Medicine Keio University Tokyo Japan
4. Division of Development and Aging Research Institute for Biomedical SciencesTokyo University of Science Noda Japan
5. Gastrointestinal Cancer Biology International Research Center for Medical Sciences (IRCMS) Kumamoto University Kumamoto Japan