1. State Key Laboratory of Forest and Soil Ecology; Institute of Applied Ecology; Chinese Academy of Sciences; Shenyang; 110164; China
2. Southwest Biological Science Center; U.S. Geological Survey; Moab; UT; 84532; USA
3. Key Laboratory of Ecosystem Network Observation and Modeling; Institute of Geographic Sciences and Natural Resources Research; Chinese Academy of Sciences; Beijing; 100101; China