1. Hokkaido Research Center Forestry and Forest Products Research Institute, Forest Research and Management Organization Sapporo Hokkaido Japan
2. Center for Biodiversity and Climate Change Forestry and Forest Products Research Institute, Forest Research and Management Organization Ibaraki Japan
3. Faculty of Life and Environmental Sciences University of Tsukuba Tsukuba Japan
4. Echigo‐Matsunoyama Museum of Natural Science Niigata Japan
5. Biological Laboratory Sapporo Campus, Hokkaido University of Education Sapporo Hokkaido Japan