1. Royal Botanic Gardens, Kew Richmond UK
2. Faculty of Science University of the Ryukyus Okinawa Japan
3. Okinawa Prefecture Environment Science Center Okinawa Japan
4. School of Environment The University of Auckland Auckland New Zealand
5. Marine and Terrestrial Field Ecology, Tropical Biosphere Research Center University of the Ryukyus Okinawa Japan
6. Institute of Oceanography National Taiwan University Taipei Taiwan
7. School of Biological Sciences and Swire Institute of Marine Science The University of Hong Kong Hong Kong China
8. Institute of Statistics National Tsing Hua University Hsin‐Chu Taiwan