1. Biodiversity Division National Institute for Environmental Studies Tsukuba Japan
2. The Kagoshima University Museum Kagoshima University Kagoshima Japan
3. Faculty of Environmental Earth Science Hokkaido University Sapporo Japan
4. Faculty of Agriculture Kagoshima University Kagoshima Japan
5. Research Center for the Pacific Islands Kagoshima University Kagoshima Japan
6. Kyushu Open University Fukuoka Japan
7. Subtropical Field Science Center, Faculty of Agriculture University of the Ryukyus Kunigami Japan