1. Graduate School of Environmental Science Hokkaido University Sapporo Japan
2. Akkeshi Marine Station, Field Science Center for Northern Biosphere Hokkaido University Akkeshi Japan
3. Marine Environmental Information Group Port and Airport Research Institute Yokosuka Japan
4. Institute of Marine Sciences, UNC Chapel Hill Morehead City North Carolina
5. Integrative Aquatic Biology, Onagawa Field Center, Graduate School of Agricultural Science Tohoku University Onagawa Japan