1. Department of Applied Biological Science Tokyo University of Science 2641 Yamazaki Noda Chiba 278–8510 Japan
2. Graduate School of Agriculture Ehime University 3‐5‐7 Tarumi Matsuyama Ehime 790‐8566 Japan
3. Graduate School of Life and Environmental Sciences Kyoto Prefectural University 1‐5 Hangi‐cho, Shimogamo, Sakyo‐ku Kyoto 606‐8522 Japan
4. Biotechnology Research Department Kyoto Prefectural Agriculture Forestry and Fisheries Technology Center Kitainayazuma Oji 74, Seika‐cho, Soraku‐gun Kyoto 619‐0244 Japan
5. Japan International Research Center for Agricultural Sciences 1‐1 Ohwashi Tsukuba Ibaraki 305‐8686 Japan
6. Department of Biological Science Graduate School of Science The University of Tokyo Bunkyo‐ku Tokyo 113‐0033 Japan