1. Department of Biology; University of Iowa; Iowa City Iowa 52242
2. Institute for Biodiversity and Ecosystem Dynamics (IBED); University of Amsterdam; P.O. Box 94248 1090GE Amsterdam The Netherlands
3. Department of Ecology and Evolution; University of Lausanne; CH-1015 Lausanne Switzerland
4. Academic Medical Center (AMC); University of Amsterdam; 1105 AZ Amsterdam The Netherlands