1. Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
2. Yunnan Key Laboratory for Wild Plant Resources Kunming Institute of Botany, Chinese Academy of Sciences Kunming 650201 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Key Laboratory of Southern Subtropical Plant Diversity Fairy Lake Botanical Garden, Shenzhen & Chinese Academy of Science Shenzhen 518004 China
5. Institute of Plant Stress Biology, State Key Laboratory of Crop Stress Adaptation and Improvement, School of Life Sciences, Henan University Kaifeng 475001 China
6. Department of Biology East Carolina University Greenville NC 27858 USA