1. College of Horticulture, South China Agricultural University/Key Laboratory of Biology and Genetic Improvement of Horticultural Crops (South China) Ministry of Agriculture and Rural Affairs/Guangdong Vegetables Engineering Research Center Guangzhou510642 GuangdongChina
2. Guangdong Provincial Key Laboratory of Utilization and Conservation of Food and Medicinal Resources in Northern Region, Henry Fok School of Biology and Agricultural Shaoguan University Shaoguan512023 GuangdongChina
3. College of Horticulture and Landscape Architecture Zhongkai University of Agriculture and Engineering Guangzhou 510225 China