1. State Key Laboratory of Vegetation and Environmental Change Institute of Botany, Chinese Academy of Sciences Beijing 100093 P.R. China
2. Jiangsu Key Laboratory of Crop Genetics and Physiology, College of Horticulture and Plant Protection Yangzhou University Yangzhou 225009 P.R. China
3. Department of Biology University of Kentucky Lexington KY 40506 USA
4. Department of Plant and Soil Sciences University of Kentucky Lexington KY 40546 USA
5. Department of Biology City University of New York Staten Island NY 10314 USA