1. State Key Laboratory of Crop Biology Shandong Agricultural University Tai'an Shandong 271018 China
2. Shandong Provincial Key Laboratory for Biology of Vegetable Diseases and Insect Pests College of Plant Protection Shandong Agricultural University Tai'an Shandong 271018 China
3. Institute of Plant Protection and Soil Fertilizer Hubei Academy of Agricultural Sciences Wuhan Hubei 430064 China
4. State Key Laboratory of Hybrid Rice College of Life Sciences Wuhan University Wuhan 430072 China
5. Hubei Hongshan Laboratory Wuhan University Wuhan Hubei 430070 China