1. State Key Laboratory of Grassland Agro‐Ecosystems, and College of Ecology Lanzhou University Lanzhou China
2. Key Laboratory of Freshwater Fish Reproduction and Development (Ministry of Education), School of Life Sciences Southwest University Chongqing China
3. Key Laboratory for Conserving Wildlife with Small Populations in Yunnan Southwest Forestry University Kunming China
4. State Key Laboratory of Genetic Resources and Evolution Kunming Institute of Zoology, Chinese Academy of Sciences Kunming China