1. School of Natural Sciences, College of Environmental Sciences and Engineering Bangor University Bangor UK
2. Biodiversity and Natural Resources Program International Institute for Applied Systems Analysis (IIASA) Laxenburg Austria
3. Royal Botanic Garden Edinburgh Edinburgh UK
4. Grantham Centre for Sustainable Futures and School of Biosciences University of Sheffield Sheffield UK
5. Ecology and Biodiversity, Department of Biology Utrecht University Utrecht The Netherlands