1. Graduate School of Life and Environmental Sciences; University of Tsukuba; Tsukuba 305-8572 Japan
2. Department of Biology; Faculty of Science; Ehime University; 2-5 Bunkyo-cho Matsuyama Ehime 790-8577 Japan
3. Department of Biology; Faculty of Science; Toyama University; 3190 Gofuku Toyama 930-8555 Japan