1. Department of Systematics; Evolution and Climate Change; Biodiversity and Climate Research Centre (BiK-F)/Senckenberg and Goethe University; Senckenberganlage 25 D-60325 Frankfurt am Main Germany
2. Department of Molecular Evolution and Systematics of Plants; Leipzig University; Johannisallee 21-23 D-04103 Leipzig Germany
3. Department of Botany; Swedish Museum of Natural History; Box 50007 SE-104 05 Stockholm Sweden
4. University of Oxford; Department of Plant Sciences; Oxford OX1 3RB UK
5. German Centre for Integrative Biodiversity Research (iDiv) Halle-Jena-Leipzig; D-04103 Leipzig Germany