1. School of Biological and Biomedical Sciences; Durham University; South Road Durham DH1 3LE UK
2. Climate Change and BioAdaptation Division; South African National Biodiversity Institute; Kirstenbosch Research Centre; P/Bag X7 Claremont 7735 Cape Town South Africa
3. School of Agricultural, Earth and Environmental Sciences; University of Kwazulu-Natal; Pietermaritzburg Campus Pvt Bag X101 Scottsville 3209 South Africa
4. Percy FitzPatrick Institute of African Ornithology; DST/NRF Centre of Excellence; University of Cape Town; Rondebosch 7701 Cape Town South Africa
5. School of Geographical Sciences; University of Bristol; University Road Bristol BS8 1SS UK