1. Frontier Research Institute for Interdisciplinary Sciences (FRIS); Tohoku University; Aoba-ku, Sendai 980-8578 Japan
2. Department of Developmental Biology and Neurosciences; Graduate School of Life Sciences; Tohoku University; Aoba-ku, Sendai 980-8578
3. Department of Zoology; Graduate School of Science; Kyoto University; Kitashirakawa, Sakyo-ku Kyoto Japan
4. AMED Core Research for Evolutional Science and Technology (AMED-CREST); Japan Agency for Medical Research and Development (AMED); Chiyoda-ku Tokyo 100-0004 Japan