1. Misaki Marine Biological Station; Graduate School of Science and Center for Marine Biology; The University of Tokyo; 1024 Koajiro Misaki Miura Kanagawa 238-0225 Japan
2. Department of Biological Sciences; Graduate School of Science; The University of Tokyo; 7-3-1 Hongo Bunkyo-ku Tokyo 113-0033 Japan
3. Institute for Amphibian Biology; Graduate School of Science; Hiroshima University; 1-3-1 Kagamiyama Higashi-Hiroshima Hiroshima 739-8526 Japan