1. Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresources, College of Tropical Crops Hainan University Haikou 570228 China
2. Hainan Yazhou Bay Seed Laboratory, College of Tropical Crops, Sanya Nanfan Research Institute Hainan University Sanya 572025 China
3. Genomics and Genetic Engineering Laboratory of Ornamental Plants, College of Agriculture and Biotechnology Zhejiang University Hangzhou 310058 China
4. Hainan Institute of Zhejiang University Sanya 572025 China